Publications Bernhard A. Huber      last update: Sept. 2016

In press

Huber BA. In press
Spider diversity and endemism in a South American hotspot: 20 new species of Carapoia (Araneae: Pholcidae) from Brazil’s Atlantic Forest. Zootaxa


Huber BA, Nuñeza OM, Leh Moi Ung C. 2016. The Philippine hair wax spiders and their relatives: Revision of the Pholcus bicornutus species group (Araneae, Pholcidae). European Journal of Taxonomy 225: 1-34.

Huber BA, Koh JKH, Ghazali ARM, Nuñeza O, Leh Moi Ung C, Petcharad B. 2016. New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae). European J Taxonomy 200: 1-45.

Huber BA, Petcharad B, Leh Moi Ung C, Koh JKH, Ghazali ARM. 2016. The Southeast Asian Pholcus halabala species group (Araneae, Pholcidae), new data from field observations and ultrastructure. European Journal of Taxonomy 190: 1-55.

Huber BA. 2016. A new genus of litter-dwelling pholcine spiders from Sarawak (Araneae, Pholcidae). European Journal of Taxonomy 186: 1-15.

Huber BA, Leh Moi Ung C. 2016. The Panjange nigrifrons group in Borneo (Araneae: Pholcidae): high diversity in Sarawak, apparent absence in Sabah. European Journal of Taxonomy 184: 1-32.

Astrin, J.J., Höfer, H., Spelda, J., Holstein, J., Bayer, S., Hendrich, L., Huber, B.A., Kielhorn, K.H., Krammer, H.J., Lemke, M., Monje, J.C., Morinière, J., Rulik, B., Petersen, M., Janssen, H., Muster, C. 2016. Towards a DNA barcode reference database for spiders and harvestmen of Germany. PLoS ONE 11(9): 1-24.


Huber BA. 2015. Small scale endemism in Brazil's Atlantic Forest: 14 new species of Mesabolivar (Araneae, Pholcidae), each known from a single locality. Zootaxa 3942: 1-60.

Huber BA, Neumann J, Rehfeldt S, Grabolle A, Reiser N. 2015. Back in Europe: Quamtana spiders (Araneae: Pholcidsae) in Germany. Arachnologische Mitteilungen 50: 51-56.

Huber BA, Nuñeza OM. 2015. Evolution of genital asymmetry, exaggerated eye stalks, and extreme palpal elongation in Panjange spiders (Araneae: Pholcidae). European Journal of Taxonomy 169: 1-46.

Huber BA, Nuñeza OM, Leh Moi Ung C. 2015. Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78.

Huber BA, Petcharad B, Bumrungsri S. 2015. Revision of the enigmatic Southeast Asian spider genus Savarna (Araneae, Pholcidae).
European Journal of Taxonomy 160: 1-23.

Huber BA. 2014. Pholcidae. In: Biodiversidad de Artrópodos Argentinos, Vol. 3 (eds. S. Roig-Juñent, L.E. Claps, J.J. Morrone), pp. 131-140. Sociedad Entomológica Argentina.
PDF (proofs)

Huber BA. 2014. Progress and prospects in taxonomy: what is our goal and are we ever going to reach it? Journal of Arachnology 42: 142-147.

Huber BA, Carvalho LS, Benjamin SP. 2014.
On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae). Invertebrate Systematics 28: 432-450.

Huber BA, Colmenares PA, Ramirez MJ. 2014. Fourteen new generic and ten new specific synonymies in Pholcidae (Araneae), and transfer of Mystes Bristowe to Filistatidae. Zootaxa 3847: 413-422.

Huber BA, Dimitrov D. 2014. Slow genital and genetic but rapid non-genital and ecological differentiation in a pair of spider species (Araneae, Pholcidae). Zoologischer Anzeiger 253: 394-403.

Huber BA, Le Gall P, Mavoungou JF. 2014. Pholcid spiders from the Lower Guinean region of Central Africa: an overview, with descriptions of seven new species (Araneae, Pholcidae). European Journal of Taxonomy 81: 1-46.


Dimitrov D, Astrin JJ, Huber BA. 2013. Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group. Cladistics 29: 132-146.
Supporting information: Figures S1-S10; Tables S1-S7.

Huber BA. 2013. Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae). Zootaxa 3713: 1-160.
PDF (27 MB)

Huber BA, Kwapong P. 2013. West African pholcid spiders: an overview, with descriptions of five new species (Araneae, Pholcidae). European Journal of Taxonomy 59: 1-44. DOI: 10.5852/ejt.2013.59

Huber BA, Pérez-González A, Astrin JJ, Blume C, Baptista R. 2013. Litoporus iguassuensis Mello-Leitão, 1918 (Araneae, Pholcidae): camouflaged retreat, sexual dimorphism, female color polymorphism, intra-specific genital variation, and description of the male. Zoologischer Anzeiger 252: 511-521. PDF;


Huber BA. 2012. Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae). Zootaxa 3461: 1-138.
PDF; (16 MB)

Huber BA, Warui CM. 2012.
East African pholcid spiders: an overview, with descriptions of eight new species (Araneae, Pholcidae). European Journal of Taxonomy 29: 1-44.
PDF; (20 MB);

Huber BA. 2011.
Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1-509.

PDF (high resolution, 206 MB)
PDF (low resolution, 36 MB)

Huber BA. 2011.
Phylogeny and classification of Pholcidae (Araneae): an update. Journal of Arachnology 39: 211-222.

Huber BA, Rheims CA. 2011. Diversity and endemism of pholcid spiders in Brazil’s Atlantic Forest, with descriptions of four new species of the Atlantic Forest endemic genus Tupigea (Araneae: Pholcidae). Journal of Natural History 45: 275-301.

Huber BA. 2010. Mating positions and the evolution of asymmetric insect genitalia. Genetica 138: 19-25. PDF;

Huber BA, Fischer N, Astrin JJ. 2010.
High level of endemism in Haiti's last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules. Zoological Journal of the Linnean Society 158: 244-299.

Huber BA, Lankhorst S. 2010. Non-insect arthropod types in the ZFMK collection, Bonn (Acari, Araneae, Scorpiones, Pantopoda, Amphipoda). Bonn zoological Bulletin 58: 217-226. PDF PDF update Sept. 2016

Eberhard WG, Huber BA. 2010.
Spider genitalia: precise maneuvers with a numb structure in a complex lock. Pp. 249-284.
In: J. L. Leonard & A. Córdoba-Aguilar (eds.) Evolution of primary sexual characters in animals. Oxford University Press, Oxford, U.K.

Huber BA. 2009.
Life on leaves: leaf-dwelling pholcids of Guinea, with emphasis on Crossopriza cylindrogaster, a spider with inverted resting position, pseudo-eyes, lampshade web, and tetrahedral egg-sac (Araneae: Pholcidae). Journal of Natural History 43: 2491-2523.

Huber BA. 2009. Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides Roewer to Filistatidae (Araneae).
Zootaxa 1970: 64-68; 1977: 68 (erratum). PDF

Huber BA, Astrin JJ. 2009. Increased sampling blurs morphological and molecular species limits: revision of the Hispaniolan endemic spider genus Tainonia (Araneae: Pholcidae). Invertebrate Systematics 23: 281-300. PDF;

Huber BA, Schütte A. 2009. Preliminary notes on leaf-dwelling Metagonia spiders (Araneae: Pholcidae) in the Esquinas Rainforest near La Gamba, Costa Rica: leaf preference, mimesis, and web structure. Contributions to Natural History (Bern) 12: 681-697.

Huber BA, Fleckenstein N. 2008. Comb-hairs on the fourth tarsi in pholcid spiders (Araneae, Pholcidae).
Journal of Arachnology 36: 232-240. PDF

Beatty JA, Berry JW, Huber BA. 2008.
The pholcid spiders of Micronesia and Polynesia (Araneae, Pholcidae).
Journal of Arachnology 36: 1-25.

Astrin JJ, Misof B, Huber BA. 2007. The pitfalls of exaggeration: molecular and morphological evidence suggests Kaliana is a synonym of Mesabolivar (Araneae: Pholcidae). Zootaxa 1646: 17-30.

Huber BA. 2007. Two new genera of small, six-eyed pholcid spiders from West Africa, and first record of Spermophorides for mainland Africa (Araneae: Pholcidae). Zootaxa 1635: 23-43.

Huber BA, El Hennawy H. 2007. On Old World ninetine spiders (Araneae: Pholcidae), with a new genus and species and the first record for Madagascar. Zootaxa 1635: 45-53.

Huber BA, Sinclair B, Schmitt M. 2007.
The evolution of asymmetric genitalia in spiders and insects. Biological Reviews 82: 647-698.

O, Brescovit AD, Candiani DF, Huber BA. 2007. Three new species of Mesabolivar (Aranea, Pholcidae) from leaf litter in urban environments in the city of São Paulo, São Paulo, Brazil. Iheringia 97: 168-176. PDF;

Machado ÉO, Yamamoto FU, Brescovit AD, Huber BA. 2007. Three new ground living pholcid species (Araneae: Pholcidae) from Parque Estadual da Cantareira, São Paulo, São Paulo, Brazil. Zootaxa 1582: 27–37.

Useta G, Huber BA, Costa FG. 2007. Spermathecal morphology and sperm dynamics in the female Schizocosa malitiosa (Araneae: Lycosidae). European Journal of Entomology 104: 777-785.

Astrin JJ, Huber BA, Misof B, Kluetsch CFC. 2006.
Molecular taxonomy in pholcid spiders (Pholcidae, Araneae): evaluations of species identification methods using CO1 and 16S rRNA.
Zoologica Scripta 35: 441-457.  PDF;

Huber BA. 2006.
Cryptic female exaggeration: the asymmetric female internal genitalia of Kaliana yuruani (Araneae: Pholcidae).
Journal of Morphology 276: 705-712.  PDF

Huber BA, Wunderlich J. 2006.
Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae). Journal of Natural History 40: 2341-2360.  PDF

Michalik P, Huber BA. 2006.
Spermiogenesis in Psilochorus simoni (Berland, 1911) (Pholcidae, Araneae): evidence for considerable within-family variation in sperm structure and development.
Zoology 109: 14-25. PDF;

Bruvo-Madaric B, Huber BA, Steinacher A, Pass G. 2005.
Phylogeny of pholcid spiders (Araneae:Pholcidae): combined analysis using morphology and molecules. Molecular Phylogenetics and Evolution 37: 661-673.  PDF

Funke S, Huber BA. 2005.
Allometry of genitalia and fighting structures in Linyphia triangularis (Araneae: Linyphiidae).
Journal of Arachnology 33: 870-872.  PDF

Huber BA. 2005.
High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae).
Zoologica 155: 1-126.  PDF (16 MB)

Huber BA. 2005.
Sexual selection research on spiders: progress and biases.
Biological Reviews 80: 363-385.  PDF;

Huber BA. 2005.
Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae).
Zoologische Mededelingen 79-2(4): 61-172.  PDF (4.3 MB)

Huber BA. 2005.
Pholcidae. In: Ubick, D., Paquin, P., Cushing, P.E., Roth, V. (eds)
Spiders of North America: an identification manual, pp. 194-196.  PDF

Huber BA. 2005.
The pholcid spiders of Africa (Araneae: Pholcidae): state of knowledge and directions for future research. In: Huber B.A., Sinclair, B.J., Lampe, K.-H. (eds.)
African Biodiversity: Molecules, Organisms, Ecosystems, pp. 181-186. Springer Verlag.  PDF

Huber BA. 2005.
Evidence for directional selection on male abdomen size in Mecolaesthus longissimus (Araneae, Pholcidae).
Journal of Arachnology 33: 573-581. PDF

Huber BA. 2005.
Revision and cladistic analysis of the spider genus Carapoia González-Sponga (Araneae: Pholcidae), with descriptions of new species from Brazil’s Atlantic forest.
Invertebrate Systematics 19: 541-556.  PDF

Huber BA, Benjamin S. 2005.
The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of Wanniyala new genus (Araneae: Pholcidae).
Journal of Natural History 39(37): 3305-3319. PDF;

Huber BA, Brescovit AD, Rheims CA. 2005.
Exaggerated female genitalia in two new spider species (Araneae: Pholcidae), with comments on genital evolution by female choice versus antagonistic coevolution.
Insect Systematics and Evolution 36: 285-292.  PDF

Huber BA, Pérez-G, A, Baptista RLC. 2005.
Leptopholcus (Araneae: Pholcidae) in continental America: rare relicts in low precipitation areas.
Bonner zoologische Beiträge 53(1/2): 99-107.  PDF (5 MB)

Huber BA, Rheims CA, Brescovit AD. 2005.
Speciation without changes in genital shape: a case study on Brazilian pholcid spiders (Araneae: Pholcidae).
Zoologischer Anzeiger 243: 273-279.

Huber BA, Rheims CA, Brescovit AD. 2005.
Two new species of litter-dwelling Metagonia spiders (Araneae, Pholcidae) document both rapid and slow genital evolution.
Acta Zoologica (Stockholm): 86: 33-40.  PDF;

Huber BA. 2004.
Evidence for functional segregation in the directionally asymmetric male genitalia of the spider Metagonia mariguitarensis (Gonzalez-Sponga) (Pholcidae: Araneae).
Journal of Zoology, London 262: 317-326.  PDF

Huber BA. 2004.
Evolutionary transformation from muscular to hydraulic movements in spider (Arachnida, Araneae) genitalia: a study based on histological serial sections.
Journal of Morphology 261: 364-376.  PDF;

Huber BA. 2004.
The significance of copulatory structures in spider systematics. In: Schult, J. (ed.) Studien zur Theorie der Biologie, Band 5, Biosemiotik - Praktische Anwendung und Konsequenzen fuer die Einzeldisziplinen, VWB-Verlag fuer Wissenschaft und Bildung, Berlin. 

Huber BA, Hopf S. 2004.
Color dimorphism in adults and juveniles of Buitinga safura (Araneae, Pholcidae).
Journal of Arachnology 32: 336-340.  PDF

Huber BA. 2003.
Cladistic analysis of Malagasy pholcid spiders reveals generic level endemism: Revision of Zatavua n. gen. and Paramicromerys Millot (Pholcidae, Araneae).
Zoological Journal of the Linnean Society 137: 261-318. PDF

Huber BA. 2003.
High species diversity in one of the dominant groups of spiders in East African montane forests (Araneae: Pholcidae: Buitinga n. gen., Spermophora Hentz).
Zoological Journal of the Linnean Society 137: 555-619. PDF (5MB)

Huber BA. 2003.
Southern African pholcid spiders: revision and cladistic analysis of Quamtana gen. nov. and Spermophora Hentz (Araneae: Pholcidae), with notes on male-female covariation.
Zoological Journal of the Linnean Society 139: 477-527.  PDF;

Huber BA. 2003.
On the terminology of polytomies.
Cladistics 19: 273.  PDF (proofs)

Huber BA. 2003.
Rapid evolution and species-specificity of arthropod genitalia: fact or artifact?
Organisms Diversity & Evolution 3: 63-71.  PDF

Huber BA, Brescovit AD. 2003.
Ibotyporanga Mello-Leitão: tropical spiders in Brazilian semi-arid habitats (Araneae: Pholcidae).
Insect Systematics and Evolution 34: 15-20. PDF;

Huber BA. 2002.
Functional morphology of the genitalia in the spider Spermophora senoculata (Pholcidae, Araneae).
Zoologischer Anzeiger 241: 105-116.  PDF (5MB)

Huber BA. 2002.
Ninetis russellsmithi n. sp., an unusual new pholcid spider species from Malawi (Araneae: Pholcidae).
Journal of Insect Science 2.4: 1-3.  PDF

Huber BA. 2001.
The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships.
Bulletin of the American Museum of Natural History 260: 1-144. PDF  (20 MB);2

Huber BA, Pérez-González A. 2001.
A new genus of pholcid spiders (Araneae: Pholcidae) endemic to Western Cuba, with a case of female genitalic dimorphism.
American Museum Novitates 3329: 1-23. PDF (11 MB);<0001:ANGOPS>2.0.CO;2

Huber BA, Pérez-González A. 2001.
Female genital dimorphism in a spider (Araneae: Pholcidae).
Journal of Zoology, London 255: 301-304. PDF

Huber BA, van Harten A. 2001.
Ninetis subtilissima Simon, 1890 (Araneae: Pholcidae): Redescription and SEM ultrastructure.
American Museum Novitates 3336: 1-7.  PDF<0001:NSSAPR>2.0.CO;2

Huber BA, Zhu M-S. 2001.
On Taczanowski’s pholcids, with three new synonymies in Pholcidae (Araneae).
Bulletin of the British arachnological Society 12(3): 151-152. PDF

Huber BA. 2000.
New World pholcid spiders (Araneae: Pholcidae): a revision at generic level.
Bulletin of the American Museum of Natural History 254: 1-348. PDF (8 MB)<0001:NWPSAP>2.0.CO;2

Eberhard WG, Huber BA, Rodriguez RL. 1999.
Don’t forget the biology: a reply to Green.
Evolution 53(5): 1624-1627. PDF;

Huber BA. 1999.
Sexual selection in pholcid spiders (Araneae, Pholcidae): artful chelicerae and forceful genitalia.
Journal of Arachnology 27(1): 135-141.  PDF

Huber BA, Deeleman-Reinhold CL, Pérez-González A. 1999.
The spider genus Crossopriza (Araneae, Pholcidae) in the New World.
American Museum Novitates 3262: 1-10. PDF;

Pérez-González A, Huber BA. 1999.
Metagonia debrasi n. sp., the first species of the genus Metagonia Simon in Cuba (Pholcidae, Araneae).
Revue Arachnologique 13(4): 69-72.  PDF

Eberhard WG, Huber BA. 1998.
Courtship, copulation, and sperm transfer in Leucauge mariana (Araneae, Tetragnathidae) with implications for higher classification.
Journal of Arachnology 26(3): 342-368.  PDF

Eberhard WG, Huber BA. 1998.
Possible links between embryology, lack of innervation, and the evolution of male genitalia in spiders.
Bulletin of the British arachnological Society 11(2): 73-80.  PDF

Eberhard WG, Huber BA. 1998.
Copulation and sperm transfer in Archisepsis flies (Diptera, Sepsidae) and the evolution of their intromittent genitalia.
Studia Dipterologica 5(2): 217-248.

Eberhard WG, Huber BA, Rodriguez RL, Briceño D, Salas I, Rodriguez V. 1998.
One size fits all? Relationships between the size of genitalia and other body parts in 20 species of insects and spiders.
Evolution 52(2): 415-431.  PDF;

Huber BA. 1998.
On the "valve" in the genitalia of female pholcids (Pholcidae, Araneae).
Bulletin of the British arachnological Society 11(2): 41-48.  PDF

Huber BA. 1998.
Notes on the neotropical spider genus Modisimus (Pholcidae, Araneae), with descriptions of thirteen new species from Costa Rica and neighboring countries.
Journal of Arachnology 26: 19-60.  PDF (4MB)

Huber BA. 1998.
Report of some pholcid spiders collected in Guatemala and Honduras (Araneae, Pholcidae).
Revue Suisse de Zoologie 105(1): 49-80.

Huber BA. 1998.
Genital mechanics in some neotropical pholcid spiders (Araneae: Pholcidae), with implications for systematics.
Journal of Zoology, London 244: 587-599.

Huber BA. 1998.
Spider reproductive behaviour: a review of Gerhardt’s work from 1911-1933, with implications for sexual selection.
Bulletin of the British arachnological Society 11(3): 81-91.  PDF

Huber BA, Pérez-González A. 1998.
Leptopholcus delicatulus (Araneae, Pholcidae) is a valid name.
Journal of Arachnology 26: 251-256.  PDF

Huber BA. 1997.
Evidence for gustatorial courtship in a haplogyne spider (Hedypsilus culicinus: Pholcidae: Araneae).
Netherlands Journal of Zoology 47(1): 95-98.  PDF

Huber BA. 1997.
On American ‘Micromerys’ and Metagonia (Araneae, Pholcidae), with notes on natural history and genital mechanics.
Zoologica Scripta 25(4): 341-363.  PDF

Huber BA. 1997.
Redescriptions of Eugène Simon’s neotropical pholcids (Araneae, Pholcidae).
Zoosystema 19(4): 573-612.  PDF

Huber BA. 1997.
The pholcid spiders of Costa Rica (Aranae: Pholcidae).
Revista de Biologia Tropical 45(4): 1583-1634.  PDF;

Huber BA, Eberhard WG. 1997.
Courtship, copulation, and genital mechanics in Physocyclus globosus (Araneae, Pholcidae).
Canadian Journal of Zoology 74: 905-918.  PDF;

Huber BA, Senglet A. 1997.
Copulation with contralateral insertion in entelegyne spiders (Araneae: Entelegynae: Tetragnathidae).
Netherlands Journal of Zoology 47(1): 99-102.  PDF;

Huber BA. 1996.
Genitalia, fluctuating asymmetry, and patterns of sexual selection in Physocyclus globosus (Araneae: Pholcidae).
Revue Suisse de Zoologie, suppl. 1996: 289-294.

Huber BA. 1996.
On the distinction between Modisimus and Hedypsilus (Araneae, Pholcidae), with notes on behavior and natural history.
Zoologica Scripta 25(3): 233-240.

Huber BA. 1995.
Genital morphology and copulatory mechanics in Anyphaena accentuata (Anyphaenidae) and Clubiona pallidula (Clubionidae: Araneae).
Journal of Zoology, London 235: 689-702. 

Huber BA. 1995.
The retrolateral tibial apophysis in spiders - shaped by sexual selection?
Zoological Journal of the Linnean Society 113: 151-163.

Huber BA. 1995.
Copulatory mechanism in Holocnemus pluchei and Pholcus opilionoides, with notes on male cheliceral apophyses and stridulatory organs in Pholcidae (Araneae).
Acta Zoologica, Stockholm 76(4): 291-300. 

Uhl G, Huber BA, Rose W. 1995.
Male pedipalp morphology and copulatory mechanism in Pholcus phalangioides (Fuesslin, 1775) (Araneae, Pholcidae).
Bulletin of the British arachnological Society 10(1): 1-9.

Huber BA. 1994.
Spermophore morphology reveals a new synapomorphy of Oecobius and Uroctea (Araneae, Oecobiidae).
Journal of Arachnology 22: 73-74.  PDF

Huber BA. 1994.
Copulatory mechanics in the funnel-web spiders Histopona torpida and Textrix denticulata (Agelenidae, Araneae).
Acta Zoologica, Stockholm 75(4): 379-384.

Huber BA. 1994.
Funktion und Evolution komplexer Kopulationsorgane. Eine Analyse von Schnittserien durch in copula schockfixierte Spinnen.
Mitt. dtsch. Ges. allg. angew. Entomol. 9: 247-250.

Huber BA. 1994.
Genital bulb muscles in entelegyne spiders.
Journal of Arachnology 22: 75-76.  PDF

Huber BA. 1994.
Genital morphology, copulatory mechanism and reproductive biology in Psilochorus simoni (Berland, 1911) (Pholcidae; Araneae).
Netherlands Journal of Zoology 44(1-2): 85-99.  PDF;

Huber BA. 1993.
Genital mechanics and sexual selection in the spider Nesticus cellulanus (Araneae: Nesticidae).
Canadian Journal of Zoology 71: 2437-2447.  PDF

Huber BA. 1993. Female choice and spider genitalia. Boll. Accad. Gioenia Sci. Nat. (Catania) 26: 209-214.  PDF

Huber KC, Haider TS, Müller MW, Huber BA, Schweyen RJ, Barth FG. 1993.
DNA sequence data indicates the polyphyly of the family Ctenidae (Araneae).
Journal of Arachnology 21: 194-201. PDF

Huber BA. 1992.
Frontal heart and arterial system in the head of isopoda.
Crustaceana 63(1): 57-69.