Latest publications on pholcid spiders (since 2019)

2025
Geci D, Ibrahimi H, Bilalli A, Musliu M. 2025. New records of spiders (Arachnida: Araneae) from the Western Balkans. Journal of Insect Biodiversity and Systematics 11: 269-289. https://doi.org/10.61186/jibs.11.2.269
Huber BA, Meng G. 2025
. Like grains of sand: Ninetis spiders on the Arabian Peninsula (Araneae: Pholcidae). Zootaxa 5563: 290-335. https://doi.org/10.11646/zootaxa.5563.1.19
Li J, He Q, Li S, Yao Z. 2025a. Filling a zoogeographical gap in China: taxonomic descriptions of six new spider species of the Pholcus phungiformes species group (Araneae, Pholcidae). ZooKeys 1232: 285-310.
https://doi.org./10.3897/zookeys.1232.145622
Li J, Li S, Zhang X, Yao Z. 2025b. Diversity survey of Pholcus spiders (Araneae, Pholcidae) from eastern Sichuan and neighboring areas, with descriptions of six new species. ZooKeys 1240: 39-64. https://doi.org./10.3897/zookeys.1240.147905
Li J, Zhang B, Li S, Yao Z. 2025c. Taxonomic notes on four closely related spider species of the genus Khorata (Araneae, Pholcidae). ZooKeys 1244: 281-291. https://doi.org./10.3897/zookeys.1244.154422
Murcia-Moreno D, Gálvez D. 2025. Introduced spiders in Panama: species distributions and new records. Biology 14, 4: 1-16. https://doi.org/10.3390/biology14010004
Sherwood D, Krammer H-J. 2025. Studies on spider nomenclature II: a synonymy and misidentification in Asian Pholcus Walckenaer, 1805 (Araneae: Pholcidae). Arachnology 20(1): 34-36.
Valdez-Mondragón A, Nolasco-Garduño S. 2025. Ixchela azteca (Araneae: Pholcidae), a widespread spider species from Central Mexico: underestimated diversity or morphological and genetic variation? Revista Mexicana de Biodiversidad 96: e965466. https://doi.org/10.22201/ib.20078706e.2025.96.5466
Wang B, Wang Y, Li S, Wu P, Yao Z. 2025. A survey of Belisana spiders (Araneae, Pholcidae) from eastern Sichuan, China. ZooKeys 1236: 219-231.
https://doi.org./10.3897/zookeys.1236.146511
Wang Z-J, Yao Y-B, Shi Y-Y, Liu K-K. 2025. A new species of the genus Khorata Huber, 2005 from Fujian Province, China (Araneae: Pholcidae). Acta Arachnologica Sinica 34: 38-42.
https://doi.org./10.3969/j.issn.1005-9628.2025.01.007

2024
Huber BA, Meng G. 2024. Old World Micropholcus spiders, with first records of acrocerid parasitoids in Pholcidae (Araneae). ZooKeys 1213: 95-182. https://doi.org./10.3897/zookeys.1213.133178
Huber BA, Meng G, Dederichs TM, Michalik P, Forman M, Král J. 2024. Castaways: the Leeward Antilles endemic spider genus Papiamenta (Araneae: Pholcidae). Invertebrate Systematics 38: IS23052. https://doi.org/10.1071/IS23052
Huber BA, Meng G, Valdez-Mondragón A. 2024. Notes on Chisosa (Araneae: Pholcidae), with the description of a new species from Mexico. Zootaxa 5419: 217-244. https://doi.org/10.11646/zootaxa.5419.2.3
Huber BA, Meng G, Cabra García J, Carvalho LS. 2024. Thriving in dry conditions: on the Neotropical spider genus Galapa (Araneae: Pholcidae). Zootaxa 5419: 301-347. https://doi.org/10.11646/zootaxa.5419.3.1
Huber BA, Szymański H, Bennett-West A. 2024. Progress or burden? Formal description of every apparently new species available in collections is neither necessary nor useful. ZooKeys 1214: 77-90. https://doi.org./10.3897/zookeys.1214.130592
Huber BA, Meng G, Král J, Ávila Herrera IM, Carvalho LS. 2024. Diamonds in the rough: Ibotyporanga (Araneae, Pholcidae) spiders in semi-arid Neotropical environments. European Journal of Taxonomy 963: 1-169. https://doi.org/10.5852/ejt.2024.963.2687
Jang CM, Kim ST. 2024. Redescription of a poorly known spider, Pholcus kwangkyosanensis Kim & Park, 2009 (Araneae: Pholcidae) from Korea. Korean Journal of Environmental Biology 42: 172-175. https://doi.org/10.11626/KJEB.2024.42.2.172
Jang CM, Bae YS, Kim ST. 2024. Two new spider species of the genus Pholcus Walckenaer, 1805 (Araneae, Pholcidae) from Korea. Korean Journal of Environmental Biology 42: 541-547. https://doi.org/10.11626/KJEB.2024.42.4.541
Lee JG, Lee JH, Choi DY, Park SJ, Baek MJ, Kim SK. 2024
. Nine new species of the genus Pholcus Walckenaer (Araneae: Pholcidae) from South Korea. Zootaxa 5432: 179-212.
https://doi.org/10.11646/zootaxa.5432.2.3
Li J, Zhang X, Yao Z. 2024. A new spider species of the Pholcus yichengicus species group (Araneae, Pholcidae) from Hebei Province, northern China. Biodiversity Data Journal 12: e141640. https://doi.org/10.3897/BDJ.12.e141640
Nekhaeva AA, Kim LV, Yeszhanov AB. 2024
. New data on the spider fauna (Arachnida: Aranei) of the East Kyzylkum Desert, Kazakhstan. Arthropoda Selecta 33: 391-405.
Nicolosi G, Pantini P, Devincenzo U, Guariento LA, Italiano V, Zanca L, Sarà M, Isaia M. 2024. New data on spiders (Arachnida, Araneae) from the islands of the Strait of Sicily (Southern Italy) with taxonomic notes on Poecilochroa loricata Kritscher, 1996 (Araneae, Gnaphosidae) and eight new records for Europe. The European Zoological Journal 91: 1009-1034.
https://doi.org/10.1080/24750263.2004.2390869
Rittinger MA, Rodríguez RL, Escalante I. 2024. Heuristic test reveals little effect of learning and maturation on early prey capture experiences in a web-building spider. Scientific Reports 14: 11736 https://doi.org/10.1038/s41598-024-61252-7
Slowik J, Schneider K, Rickman T. 2024. Description of the female of Psilochorus inyo Slowik, 2009 (Araneae: Pholcidae) from California, with notes on its distribution and habitat. The Pan-Pacific Entomologist 100: 154-157.
Wang B, Yao Z, Zhang X. 2024. A new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from Guizhou Province, south-western China. Biodiversity Data Journal 12,  e125111: 1-12.
https://doi.org/10.3897/BDJ.12.e125111
Wang B, Li J, Li S, Yao Z. 2024. Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China. ZooKeys 1216: 265-302.
https://doi.org./10.3897/zookeys.1216.132561
Wunderlich J. 2024. New and already known spiders of seven families of the Algarve, Portugal (Araneida: Araneae). Beitr. Araneol. 17: 7-51.
Yang L, Fu C, Zhang Y, He Q, Yao Z. 2024. A survey of Pholcus spiders (Araneae, Pholcidae) from the Qinling Mountains of central China, with descriptions of seven new species. Zoosystematics and Evolution 100: 199-221. https://doi.org/10.3897/zse.100.116759
Zhang L, Li S, Yao Z. 2024. A new species of the spider genus Khorata Huber, 2005 (Araneae, Pholcidae), with a list of Khorata species from Vietnam. Biodiversity Data Journal 12: e128884. https://doi.org/10.3897/BDJ.12.e128884
Zhang L, Wang B, He Q, Yao Z. 2024. A new species of the Pholcus phungiformes species group (Araneae, Pholcidae) from Liaoning, China, with identification keys to four closely related species. ZooKeys 1193: 171-179. https://doi.org./10.3897/zookeys.1193.115640
Zhang L, Wang B, Li S, Zhang X, Yao Z. 2024. Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China. ZooKeys 1209: 315-330. https://doi.org./10.3897/zookeys.1209.127951
Zhang L, Wu Z, Li S, Yao Z. 2024. Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China. ZooKeys 1202: 255-286. https://doi.org/10.3897/zookeys.1202.121633
Zhou G, Lu J, Deng Y. 2024. A new species of Khorata Huber, 2005 (Araneae, Pholcidae) from Jiangxi Qiyunshan National Nature Reserve, southern China. Biodiversity Data Journal 12: e141018.
https://doi.org/10.3897/BDJ.12.e141018

2023
Fernández-Pérez J, Prieto CE. 2023. Nuevos datos de Araneae (excl. Troglohyphantes) para las cuevas Vasco-Cantábricas. Revista Ibérica de Aracnología 42: 137-158.
Huber BA, Meng G, Dupérré N, Astrin J, Herrera M. 2023
. Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae). European Journal of Taxonomy 909:1-63.
https://doi.org/10.5852/ejt.2023.909.2351
Huber BA. 2023. Do pholcid spiders balloon? Arachnology 19(6): 885-887.
Huber BA, Meng G, Král J, Ávila Herrera IM, Izquierdo MA. 2023. Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900: 32-80. https://doi.org/10.5852/ejt.2023.880.2173
Huber BA, Meng G, Clark HL, Cazanove G. 2023. First blind daddy long-legs spiders from Australia and Réunion (Araneae, Pholcidae). Subterranean Biology 46: 1-19. https://doi.org/10.3897/subtbiol.46.105798
Huber BA, Meng G, A Váldez-Mondragón, Král J, Ávila Herrera IM, Carvalho LS. 2023. Short-legged daddy-long-leg spiders in North America: the genera Pholcophora and Tolteca (Araneae, Pholcidae). European Journal of Taxonomy 880: 1-89. https://doi.org/10.5852/ejt.2023.880.2173
Huber BA, Meng G, Dupérré N, Herrera M, Inclán D, Wipfler B. 2023. Humpback spiders from Ecuador: relationships, prosoma ‘inflation’, and genital asymmetry (Araneae: Pholcidae: Mecolaesthus). Invertebrate Systematics 37: 117-151. https://doi.org/10.1071/IS22052
Huber BA, Meng G,
Král J, Ávila Herrera IM, Izquierdo MA, Carvalho LS. 2023. High and dry: integrative taxonomy of the Andean spider genus Nerudia (Araneae: Pholcidae). Zoological Journal of the Linnean Society 198: 534-591. https://doi.org/10.1093/zoolinnean/zlac100
Huber BA, Meng G. 2023. On the mysterious Seychellois endemic spider genus Cenemus (Araneae, Pholcidae). Arthropod Systematics and Phylogeny 81: 179-200. https://doi.org/10.3897/asp.81.e86793
Izquierdo MA, Dederichs TM, Cargnelutti F, Michalik P. 2023. Copulatory behaviour and genital mechanics suggest sperm allocation by a non-intromittent sclerite in a pholcid spider. Royal Society Open Science 10: 230263. http://doi.org/10.1098/rsos.230263
Jang C-M, Yoo J-S, Kim S-T, Bae Y-S. 2023a. Rocky area inhabiting daddy long-legs spiders, Pholcus Walckenaer, 1805 (Araneae: Pholcidae) in mountainous mixed forests from South Korea. Forests 14 (538). 1-8. https://doi.org/10.3390/f14030538
Jang CM, Bae YS, Lee SY, Yoo JS, Kim ST 2023
b. Five new species of the Pholcus phungiformes species group (Araneae, Pholcidae) from South Korea. ZooKeys 1178: 97-114.
https://doi.org/10.3897/zookeys.1178.104780
Lecigne S, Moutaouakil S, Lips J. 2023. Contribution to the knowledge of the spider fauna of Morocco (Arachnida: Araneae) - First note - On new records of cave spiders. Arachnologische Mitteilungen 66: 44-71.
Yang L, Zhao F, He Q, Yao Z. 2023. A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China. ZooKeys 1186: 175-184. https://doi.org./10.3897/zookeys.1186.105736
Zhao F, Jiang T, Yang L, He Q, Zheng G, Yao Z. 2023a. Pholcid spiders of the Pholcus phungiformes species-group (Araneae, Pholcidae) from Liaoning Province, China: an overview, with description of a new species. ZooKeys 1156: 1-14. https://doi.org/10.3897/zookeys.1156.98331
Zhao F, Yang L, Zou Q, Ali A, Li S, Yao Z. 2023b. Diversity of Pholcus spiders (Araneae: Pholcidae) in China's Lüliang Mountains: an integrated morphological and molecular approach. Insects 14(364): 1-34. https://doi.org/10.3390/insects14040364
Zhao F, Yang L, Li S, Zheng G, Yao Z. 2023c.  A further study on the Belisana spiders (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China. Zootaxa 5351: 543-558. https://doi.org/10.11646/zootaxa.5351.5.3

2022
Alioua Y, Bosmans R. 2022. First record of the genus Nita Huber & El-Hennawy, 2007 (Araneae: Pholcidae) from Algeria. Serket 18: 282-286.
Carballo-Morales JD, Delgado-Montes C, Auccacusi-Choque LV, Reyes-Ortiz M, Aisenberg A. 2022
. Niche partitioning and intraspecific shared webs in two species of Modisimus Simon, 1893 (Pholcidae: Araneae). Journal of Arachnology 50: 43-46. https://doi.og/10.1636/JoA-S-21-020
Carvalho LS, Huber BA. 2022. A replacement name for Pinocchio Huber & Carvalho, 2019 (Araneae: Pholcidae). Zootaxa 5162: 97-98. https://doi.org/10.11646/zootaxa.5162.1.7
Ceccolini F, Cianferoni F. 2022. Nomenclatural changes for two spider genera as a consequence of homonyny (Arachnida: Araneae). Arachnology 19: 14-15. https://doi.org/10.13156/arac.2022.19.1.14
Chu C, Yao Z, Wongprom P, Li S. 2022. Changminia gen. nov., a new genus of daddy-long-leg spiders (Araneae: Pholcidae) from karst caves in Southeast Asia. Zootaxa 5092: 238-246. https://doi.org/10.11646/zootaxa.5092.2.6
Cortés-Fossati F, Martín-Rodríguez I. 2022. El endemismo Holocnemus hispanicus Wiehle, 1933 (Araneae, Pholcidae) presente en la comunidad de Aragón (España). Revista Iberica de Aracnologia 41: 134.
Dederichs TM, Huber BA, Michalik P. 2022
. Evolutionary morphology of sperm in pholcid spiders (Pholcidae, Synspermiata). BMC Zoology 7: 52. https://doi.org/10.1186/s40850-022-00148-3

Geci D, Ibrahimi H, Bilalli A, Musliu M, Grapci-Kotori L, Gashi A. 2022. The spider fauna (Arachnida, Araneae) of abandoned military bunkers in Albania. Nat. Croat. 31: 71-78.
Huber BA. 2022
. Revisions of Holocnemus and Crossopriza: the spotted-leg clade of Smeringopinae (Araneae, Pholcidae). European Journal of Taxonomy 795: 1-241. https://doi.org/10.5852/ejt.2022.795.1663
Huber BA, Acurio AE. 2022. Are introduced spiders displacing native species on Galápagos? Observations on pholcid spiders (Araneae: Pholcidae). Arachnology 19: 191-198.
Huber BA, Meng G, Acurio AE, Astrin JJ, Inclán DJ, Izquierdo M, Valdez-Mondragón A. 2022. Metagonia spiders of Galápagos: blind cave-dwellers and their epigean relatives (Araneae, Pholcidae). Invertebrate Systematics 36(7): 647-678. https://doi.org/10.1071/IS21082
Král J, Ávila Herrera IM, Šťáhlavský F, Sadílek D, Pavelka J, Chatzaki M, Huber BA. 2022. Karyotype differentiation and meiosis in European clades of the spider genus Pholcus (Araneae, Pholcidae). Comparative Cytogenetics 16(4): 185-209. https://doi.org/10.3897/compcytogen.v16.i4.85059
Lu Y, Yao Z, He Q. 2022. A new species of Pholcus yichengicus species-group (Araneae, Pholcidae) from Hebei Province, China. Biodiversity Data Journal 10: e81800. https://doi.org/10.3897/BDJ.10.e81800
Lu Y, Chu C, Zhang X-Q, Li S, Yao Z-Y. 2022. Europe vs. China: Pholcus (Araneae, Pholcidae) from Yanshang-Taihang Mountains confirms uneven distribution of spiders in Eurasia. Zoological Research 43(3): 532-534. https://doi.org/10.24272/j.issn.2095-8137.2022.103
Nolasco GS, Valdez-Mondragón A. 2022. Four new species of the spider genus Physocyclus Simon, 1893 (Araneae: Pholcidae) from Mexico, with updated taxonomic identification keys. European Journal of Taxonomy 813: 173-206. https://doi.org/10.5852/ejt.2022.813.1739
Nolasco S, Valdez-Mondragón A. 2022
. To be or not to be... Integrative taxonomy and species delimitation in the daddy long-legs spiders of the genus Physocyclus (Araneae, Pholcidae) using DNA barcoding and morphology. ZooKeys 1135: 93-118.
https://doi.org/10.3897/zookeys.1135.94628

2021
Ahmed HSK 2021. Biological studies on Holocnemus pluchei (Pholcidae) when fed on various prey species. Journal of Plant Protection and Pathology 12: 163-166.
Ávila Herrera IM, Král J, Pastuchová M, Forman M, Musilová J, Kořínková T, Štáhlavský F, Zrzavá M, Nguyen P, Just P, Haddad CR, Hiřman M, Koubová M, Sadílek D, Huber BA. 2021. Evolutionary pattern of karyotypes and meiosis in pholcid spiders (Araneae: Pholcidae): implications for reconstructing chromosome evolution of araneomorph spiders. BMC Ecology and Evolution 21. 75 https://doi.org/10.1186/s12862-01750-8
Bloem GJ, Noordijk J 2021. Arachnids (Arachnida: Araneae, Mesostigmata, Pseudoscorpiones) from tropical greenhouses at Rotterdam Zoo (the Netherlands), including a pholcid spider new to Europe. Arachnologische Mitteilungen 61: 36-44.
Huber BA. 2021a. Beyond size: sexual dimorphisms in pholcid spiders. Arachnology 18: 656-677. https://doi.org/10.13156/arac.2020.18.7.656
Huber BA. 2021b. First Northwest African species of the spider genus Artema, from caves in Morocco, with notes on body size in pholcid spiders (Araneae, Pholcidae). Zootaxa 4984: 324-334. https://doi.org/10.11646/zootaxa.4984.1.23
Huber BA, Eberle J. 2021. Mining a photo library: eggs and egg sacs in a major spider family. Invertebrate Biology 140: e12349, 1-13. https://doi.org/10.1111/ivb.12349
Huber BA, Pavlek M, Komnenov M. 2021. Revision of the spider genus Stygopholcus (Araneae, Pholcidae), endemic to the Balkan Peninsula. European Journal of Taxonomy 752: 1-60. https://doi.org/10.5852/ejt.2021.752.1391
Kumada K 2021. New records of Holocnemus pluchei (Scopoli 1763) (Pholcidae) from Japan. Acta Arachnologica 70(1): 17-19. [in Japanese]
Lan T, Jäger P, Zhu W, Li S 2021. Pholcid spiders (Araneae: Pholcidae) from Southeast Asia, with descriptions of five new species. Zootaxa 4963: 545-562.
Lee JG, Lee JH, Choi DY, Park SJ, Kim AY, Kim SK 2021a
. Four new species of the genus Pholcus Walckenaer (Araneae, Pholcidae) from Korea. Journal of Species Research 10: 86-98
.
Lee JG, Lee JH, Choi DY, Park SJ, Kim AY, Kim SK 2021b. Five new species of the genus Pholcus Walckenaer (Araneae, Pholcidae) from South Korea. Zootaxa 5052: 61-77. https://doi.org/10.11646/zootaxa.5052.1.3
86-98.
Orozco-Gil M, Desales-Lara MA. 2021
. Las arañas (Arachnida: Araneae) del estado de Hidalgo, México: contribución al conocimiento de su biodiversidad. Acta Zoológica Mexicana (nueva serie) 37: 1-23.
Osorio Moreno CM 2021. Contribución al conocimiento taxonómico de arañas neotropicales del género Mecolaesthus Simon, 1893 (Araneae: Pholcidae) en Colombia. Unpublished thesis, Universidad del Atlántico, Puerto Colombia, Atlántico, Colombia. 68 pages.
Pinzon J, Kent K, Bennett R 2021. First record of Pholcus opilionoides (Schrank) (Araneae: Pholcidae) in Canada, with notes on its biology. Journal of the Entomological Society of British Columbia 118: 53-56.
Sheng H, Xu X, Yin H, Irfan M 2021. A new species of spider genus Khorata Huber, 2005 from China (Araneae: Pholcidae). Pakistan Journal of Zoology 53: 2463-2466.
Vishnudas EH, Sudhikumar AV 2021. First report of the small daddy long leg spider Micropholcus fauroti (Simon, 1887) (Araneae: Pholcidae) female from India with redescription of the male. Serket 18(1): 59-63.
Yao Z, Luo Y, Li S 2021. Tangguoa gen. nov., one new genus of daddy-long-leg spiders (Araneae: Pholcidae) from southern China. Zootaxa 4938: 131-140.
Yao Z, Wang X, Li S 2021. Tip of the iceberg: species diversity of Pholcus spiders (Araneae, Pholcidae) in the Changbai Mountains, Northest China. Zool. Res. 42: 267-271. Supplement S1-S60.
Zhu W, Li S 2021. Two new spiders of the genus Belisana (Araneae: Pholcidae) from Xishuangbanna Rainforest, China. Zootaxa 4933: 557-566.
Zhu W, Li S. 2021. Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia. Zootaxa 4963: 115-137.

2020
Breitling R 2020. South European spiders from the Duffey collection in the Manchester Museum (Arachnida: Araneae). Arachnology 18: 333-362.
Cargnelutti F, Calbacho-Rosa L, Uñates D, Costa-Schmidt LE, Córdoba-Aguilar A, Peretti AV. 2020. Copulatory behaviour increases sperm viability in female spiders. Biological Journal of the Linnean Society 131: 536-546.
García-Villafuerte MA, Valdez-Mondragón A 2020. The oldest fossils from the spider subfamily Modisiminae from the Americas: description of a new species of the genus Modisimus Simon (Araneae: Pholcidae) from the amber of Mexico with a checklist of the extant Mexican species. Journal of South American Earth Sciences 103: 102702, p. 1-14.
Huber BA 2020. Revision of the spider genus Hoplopholcus Kulczyński (Araneae, Pholcidae). Zootaxa 4726: 1-94
.
Huber BA, Villarreal O 2020. On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy 718: 1-317.
Jiménez ML, Palacios-Cardiel C, Chávez-Lara ID 2020. Las arañas (Arachnida: Araneae) de los Médanos de Samalayuca, Chihuahua y nuevos registros de especies para México. Acta Zoologica Mexicana 36: 1-23.
Lan T, Yao Z, Ali A, Zheng G, Li S 2020a. Two new species of the spider genus Pholcus Walckenaer, 1805 (Araneae: Pholcidae) from Pakistan. Zootaxa 4820: 177-185.
Lan T, Yao Z, Zheng G, Li S 2020b. Two new species of the spider genus Pholcus Walckenaer, 1805 (Araneae: Pholcidae) from Guizhou and Guangxi, China. Zootaxa 4758: 360-368.
Lan T, Yao Z, Zheng G, Wongprom P, Li S 2020c. Five new species of the spider genus Savarna Huber, 2005 (Araneae: Pholcidae) from Thailand. Zootaxa 4778: 311-328.
Nolasco S, Valdez-Mondragón A 2020. On the daddy long-legs spiders of the genus Physocyclus (Araneae: Pholcidae) from Mexico: description of a new species from Baja California Peninsula. Revista Mexicana de Biodiversidad 91: e913316.
Valdez-Mondragón A 2020. COI mtDNA barcoding and morphology for species delimitation in the spider genus Ixchela Huber (Araneae: Pholcidae), with description of two new species from Mexico. Zootaxa 4747: 54-76.
Víquez C 2020. Aracnofauna (Arachnida) de la Isla del Coco, Costa Rica, con la descripción de tres nuevas especies. Revista de Biología Tropical 68: S115-S143.
Wang X, Shaheen S, He Q, Yao Z 2020. Notes on two closely related spider species of the Pholcus phungiformes species group (Araneae, Pholcidae) from Beijing, China. ZooKeys 965: 1-16.
Xin Y, Yao Z, Li S 2020. Two new species of the spider genus Wanniyala Huber & Benjamin, 2005 (Araneae: Pholcidae) from Sri Lanka. Zootaxa 4759: 566-574.
Xu H, Zheng G, Yao Z. 2020. Two new species of the spider genus Khorata Huber, 2005 (Araneae: Pholcidae) from southern China. Zootaxa 4747: 156-166.
https://doi.org/10.11646/zootaxa.4747.1.6
Zhu W, Yao Z, Zheng G, Li S 2020a. Three new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China. Zootaxa 4742: 332-342.
Zhu W, Yao Z, Zheng G, Li S 2020b. Six new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from southern China. Zootaxa 4810: 175-197.
Zhu W, Yao Z, Zheng G, Li S 2020c. The Belisana spiders (Araneae: Pholcidae) from Tibet. Zootaxa 4802: 111-128.

2019

Armas LF de, Pérez-González A 2019. Physocyclus globosus: cannibalismo no sexual y depredación de inmaduros de Phormictopus cubensis (Araneae: Pholcidae, Theraphosidae). Reviste Ibérica de Aracnología 34: 149-150.
Baker IM, Ali HB, Fadhil HY 2019. First record of the cellar spider genus Nita Huber & El-Hennawy, 2007 (Araneae, Pholcidae) from Iraq. Bull. Iraq nat. Hist. Mus. 15: 403-411.
Barrientos JA, Bra
ñas N, Mederos J 2019. Dos especies nuevas de Pholcidae (Araneae) de Marruecos. Revista Ibérica de Aracnología 34: 3-12.
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