Literature
on
Pholcidae
latest update: January 2025

- Absolon K, Kratochvil J 1932. Zur
Kenntnis der höhlenbewohnenden Araneae der illyrischen
Karstgebiete. Mitt. Höhl. Karstforsch. 1932: 73-81.
- Ackermann G 2012. Lepidodactylus lugubris (Squamata:
Gekkonidae) als Beute von Pholcus
phalangioides (Araneae: Pholcidae). Arachnol. Mitt. 44: 14-16.
- Agnew CW, Dean DA, Smith JW
1985.
Spiders collected from peanuts and non-agricultural habitats in the
Texas
West Cross-Timbers. Southw. Naturalist 30(1): 1-12.
- Aharon S, Huber BA, Gavish-Regev E. 2017. Daddy-long-leg
giants: revision of the spider genus Artema
Walckenaer, 1837 (Araneae, Pholcidae). European J
Taxonomy 376: 1-57. http://doi.org/10.5852/ejt.2017.376
- Ahmad
NFR, Abou-Setta MM 2017. Life history of the pholcid spider, Micropholcus fauroti (Simon, 1887)
(Araneae: Pholcidae) in Egypt. Acarines 11: 31-35.
- Ahmed HSK 2021. Biological studies
on Holocnemus pluchei
(Pholcidae) when fed on various prey species. Journal of Plant
Protection and Pathology 12: 163-166.
- Aisenberg A 2022. Niche partitioning
and intraspecific
shared webs in two species of Modisimus
Simon, 1893 (Pholcidae: Araneae). Journal of Arachnology
50: 43-46. https://doi.og/10.1636/JoA-S-21-020
- Alayo D 1957. Lista de los
Arácnidos
de Cuba. Santiago de Cuba. Pp. 1-41; pl. 1-11.
- Alayon Garcia G 1994. Lista de las
arañas
(Arachnida: Araneae) de Cuba. Avacient 10: 3-29.
- Alayon Garcia G 1995. Lista de las
arañas
(Arachnida: Araneae) de Cuba. Cocuyo 4: 16-19.
- Alioua Y, Bosmans R. 2022. First
record of the genus Nita Huber
& El-Hennawy, 2007 (Araneae: Pholcidae) from Algeria. Serket 18:
282-286.
- Al-Khazali AM, Najim SA. 2018. First
records of Pholcidae (Arachnida, Araneae) from Iraq. Bull. Iraq nat.
Hist. Mus. 15(2): 179-187.
- Armas LF de, Pérez-González
A. 2019. Physocyclus globosus:
cannibalismo no sexual y depredación de inmaduros de Phormictopus cubensis (Araneae:
Pholcidae, Theraphosidae). Reviste Ibérica de Aracnología
34: 149-150.
- Astrin JJ, Huber BA, Misof B,
Klütsch
CFC 2006. Molecular taxonomy in pholcid spiders (Pholcidae,
Araneae):
An evaluation of methods using 16S rRNA and CO1. Zoologica Scripta 35:
441-457.
- Astrin JJ,
Misof B,
Huber
BA 2007. The pitfalls of exaggeration:
molecular and
morphological
evidence suggests Kaliana is a synonym of Mesabolivar
(Araneae:
Pholcidae). Zootaxa 1646: 17-30.
- Astrin
JJ, Höfer H, Spelda J, Holstein J, Bayer S, Hendrich L, Huber BA,
Kielhorn KH, Krammer HJ, Lemke M, Monje JC, Morinière J, Rulik
B, Petersen M, Janssen H, Muster C 2016. Towards a DNA barcode
reference database for spiders and harvestmen of Germany. PLoS ONE
11(9): 1-24. http://dx.doi.org/10.1371/journal.pone.0162624
- Avalos
G, Rubio GD, Bar ME, Damborsky MP 2006. Lista preliminar de la
araneofauna
(Arachnida: Araneae) del Centro-Norte de la Provincia de Corrientes,
Argentina.
Rev. Iberica Arachnol. 13: 189-194.
- Ávila Herrera IM, Král J,
Pastuchová M, Forman M, Musilová J, Kořínková T, Štáhlavský F, Zrzavá
M, Nguyen P,
Just P, Haddad CR, Hiřman M, Koubová M, Sadílek D, Huber
BA. 2021. Evolutionary pattern of karyotypes and meiosis in
pholcid spiders (Araneae:
Pholcidae): implications for reconstructing chromosome evolution of
araneomorph spiders. BMC Ecology and
Evolution 21. 75 https://doi.org/10.1186/s12862-01750-8
- Badcock HD & Oxon MA 1932.
Arachnida
from the Paraguayan Chaco. J. Linn. Soc. London 38: 1-48.
- Baert L 2013. Description of
several unknown Galápagos male and female spiders. Bull. Soc.
roy. belge Entomol. 149: 64-73.
- Baert L 2014. New spider species
(Araneae) from the Galápagos Islands (Ecuador). Bulletin de la
Société royale belge d'Entomologie 149: 263-271.
- Baert L, Maelfait J-P 1986. A
contribution
to the knowledge of the spider fauna of Galápagos (Ecuador).
Bull.
Inst. Royal Sci. Nat. Belgique, Entomol. 56: 93-123.
- Baert L, Desender K, Maelfait J-P 1991.
Spider communities of Isla Santa Cruz (Galápagos, Ecuador). J.
Biogeogr.
18: 333-340.
- Baert L, Maelfait J-P, Desender K 1989.
Results of the Belgian 1986-expedition: Araneae, and provisional
checklist
of the spiders of the Galápagos archipelago. Bull. Inst. Royal
Sci.
Nat. Belgique, Entomol. 58: 29-54.
- Baert L, Maelfait J-P, Desender K 1989.
Results of the Belgian 1988-expedition to the Galápagos islands:
Araneae.
Bull. Inst. Royal Sci. Nat. Belgique, Entomol. 59: 5-22.
- Baert L, Maelfait J-P, Hendrickx F, Desender K
2008.
Distribution and habitat preference of the spiders (Araneae) of
Galápagos. Bull. Inst. Royal
Sci.
Nat. Belgique, Entomol. 78: 39-111.
- Baker IM, Ali HB, Fadhil HY 2019.
First record of the cellar spider genus Nita Huber & El-Hennawy, 2007
(Araneae, Pholcidae) from Iraq. Bull. Iraq nat. Hist. Mus. 15: 403-411.
- Banks N 1896. New North American
spiders
and mites. Trans. Amer. ent. Soc. 23: 57-77.
- Banks N 1898. Arachnida from Baja
California
and other parts of Mexico. Proc. California Acad. Sci. (3)1(7): 205-309.
- Banks N 1902. Arachnida. In: Papers
from
the Hopkins Stanford Galapagos expedition, 1898-1899. VII.
Entomological
results (6). Proc. Wash. Acad. Sci. 4: 49-86.
- Banks N 1905. Description of new
American
spiders. Proc. Ent. Soc. Wash. 7(2-3): 94-101.
- Banks N 1913. Notes on the types of
some
American spiders in European collections. Proc. Acad. Nat. Sci. Phil.
(1913):
177-188; pl. 11-13.
- Banks N 1914. New West Indian spiders.
Bull.
Amer. Mus. Nat. Hist. 33: 639-643.
- Banks N 1921. New Californian spiders.
Proc.
California Acad. Sci. 11(7): 99-102.
- Banks N 1929. Spiders from Panama.
Bull.
Mus. comp. Zool. 69(3): 51-96; pl. 1-4.
- Banks N 1931. Arachnida. In: The
Norwegian
zoological expedition to the Galapagos Islands 1925, conducted by Alf
Wollebaek.
Nyt. Mag. Naturv. 68: 271-278; pl. 1-2.
- Banks N, Newport NM, Bird RD
1932.
Oklahoma spiders. Publ. Univ. Oklahoma 4: 7-49.
- Barrion AT, Litsinger
JA 1995.
Riceland Spiders of South and Southeast Asia. CAB International,
Wallingford,
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- Barrientos JA, Brañas N, Mederos J.
2019. Dos especies nuevas de Pholcidae (Araneae) de Marruecos.
Revista Ibérica de Aracnología 34: 3-12.
- Barrientos JA, Ferrández MA 1982.
La colección de Araneidos del Departamento de Zoología de
la Universidad de Salamanca, III: arañas migalomorfas,
haploginas, y cribeladas. Bol. Asoc. esp. Entom. 5: 75-86.
- Barrientos JA,
Bosco Febrer J 2017. Arañas (Arachnida, Araneae) de
Menorca (Islas Baleares, España). Nuevos Datos. Rvista
Ibérica de Aracnología 31: 8-24.
- Barriga JC, Jiménez-Valverde A,
Morano E, Moreno AG, Melic A 2006.
Aranas de la provincia de Ciudad Real (Arachnida: Araneae) (Castilla la
Mancha, Espana). Rev. Ibérica de Aracnología 13: 125-142.
- Barros Machado A de 1944. Observations
inédites sur le colulus et les filières de quelques
Aranéides,
accomagnées de notes sur la morphologie comparée de
filières.
Arquivos do Museu Bocage ...:
- Bauer T, Wendt I, Holstein J, Gabriel G.
2016. Crossopriza lyoni new
to Germany (Araneae: Pholcidae). Arachnol. Mitt. 52: 4-6.
- Bayram A, Allahverdi H,
Danisman T, Yigit N, Kunt KB 2008. A
new genus and species record
from Turkey: Psilochorus simoni
(Berland, 1911) (Araneae, Pholcidae). Turk. J. Arach. 1: 91-97.
- Beatty JA, Berry JW,
Huber BA 2008. The
pholcid spiders of Micronesia and Polynesia (Araneae, Pholcidae). J.
Arachnol. 36: 1-25.
- Becker L 1896. Les arachnides de
Belgique.
Ann. Mus. Roy. Hist. nat. Belg. 12: 1-3788.
- Bedot M 1909. Sur la faune de
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- Benhadi-Marín
J 2010. First records of Psilochorus
simoni (Berland, 1911) (Araneae: Pholcidae) in the Iberian
Peninsula. Revista Ibérica de Aracnología 18: 101-104.
- Benhadi-Marín
J, Barrientos JA, Ferrández MA, Henruqies S, Morano E
2013. Second annotated list of selected spider families
(Arachnida: Araneae) held at the University of León (Spain).
Revista Ibérica de Aracnología 22: 71-78.
- Bennett R. 2014. COSEWIC assessment
and status report on the northwestern cellar spider Psilochorus hesperus in Canada.
Committee on the Status of Endagered Wildlife in Canada. Ottawa. X + 42
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- Berland L 1911. Sur deux
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- Berland L 1919. Diagnoses
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- Berland L 1924. Araignées de
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- Berland L 1938. Araignées des
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AD 2003. Ibotyporanga Mello-Leitão: tropical
spiders
in Brazilian semi-arid habitats (Araneae: Pholcidae). Ins. Syst. Evol.
34:
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the gaps: descriptions of unnamed species included in the latest
moelcular phylogeny of Pholcidae (Araneae). Zootaxa 4546: 1-96. https://doi.org/10.1164/zootaxa.4546.1.1
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global species richness from megatransect data and
undetected species estimates.
Contributions to
Zoology 88: 42-53. https://doi.org/10.1163/18759866-20191347
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Slow genital and genetic but rapid non-genital and ecological
differentiation in a pair of spider species (Araneae, Pholcidae). Zoologischer
Anzeiger 253: 394-403.
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Courtship,
copulation, and genital mechanics in Physocyclus globosus
(Araneae,
Pholcidae). Can. J. Zool. 74: 905-918. http://dx.doi.org/10.1139/z97-109
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Mining a photo library: eggs and egg sacs in a major spider family.
Invertebrate Biology 140: e12349, 1-13. https://doi.org/10.1111/ivb.12349
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Hennawy H 2007.
On Old World ninetine spiders
(Araneae: Pholcidae), with
a
new
genus and species and the first record for Madagascar. Zootaxa 1635: 45-53.
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BA, Fleckenstein N 2008. Comb-hairs
on the fourth tarsi in pholcid spiders (Araneae, Pholcidae). J. Arachnol. 36: 232-240. http://dx.doi.org/10.1636/CSh07-71.1
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S 2004. Color dimorphism in adults and juveniles of Buitinga
safura
(Araneae, Pholcidae). J. Arachnol. 32: 336-340.
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African pholcid spiders: an overview, with descriptions of five new
species (Araneae, Pholcidae).
European J. Taxonomy 59: 1-44.
- Huber BA, Meng G. 2023. On the
mysterious Seychellois endemic spider genus Cenemus (Araneae, Pholcidae). Arthropod Systematics and Phylogeny 81:
179-200. https://doi.org/10.3897/asp.81.e86793
- Huber BA, Meng G. 2024. Old World Micropholcus spiders, with first
records of acrocerid parasitoids in Pholcidae (Araneae). ZooKeys 1213: 95-182.
https://doi.org./10.3897/zookeys.1213.133178
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BA, Meng G. 2025. Like grains of
sand: Ninetis
spiders
on the Arabian Peninsula (Araneae: Pholcidae). Zootaxa
5563:
290-335. https://doi.org/10.11646/zootaxa.5563.1.19
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BA Nuñeza OM 2015. Evolution of genital
asymmetry, exaggerated eye stalks,
and extreme
palpal elongation in Panjange spiders
(Araneae: Pholcidae). European
Journal of Taxonomy 169: 1-46. http://dx.doi.org/10.5852/ejt.2015.169
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Leptopholcus
delicatulus (Araneae, Pholcidae) is a valid
name.
J. Arachnol. 26: 251-256.
- Huber BA, Pérez González A 2001a. A
new genus of pholcid spiders (Araneae: Pholcidae) endemic to Western
Cuba,
with a case of female genitalic dimorphism. Amer. Mus. Novitates 3329:
1-23.
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genital dimorphism in a spider (Araneae: Pholcidae). J. Zool. London.
255:
301-304. http://dx.doi.org/10.1017/S095283690100139X
- Huber BA, Rheims CA 2011. Diversity
and endemism of pholcid spiders in
Brazil’s Atlantic
Forest, with descriptions of four new species of the Atlantic Forest
endemic
genus Tupigea (Araneae: Pholcidae). J. Nat. Hist. 45: 275-301.
- Huber BA, Schütte A 2009. Preliminary notes
on leaf-dwelling Metagonia spiders (Araneae:
Pholcidae) in the Esquinas Rainforest near La Gamba, Costa Rica: leaf
preference, mimesis, and web structure. Contr.
Nat. Hist. (Bern) 12: 681-697.
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subtilissima Simon, 1890 (Araneae: Pholcidae): Redescription and
SEM
ultrastructure. Amer. Mus. Novitates 3336: 1-7.
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Venezuelan pholcid spiders
(Araneae, Pholcidae). European Journal of Taxonomy 718: 1-317. https://doi.org/10.5852/ejt.2020.718.1101
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African pholcid spiders: an overview, with descriptions of eight new
species (Araneae, Pholcidae). European
Journal of
Taxonomy 29: 1-44.
- Huber
BA, Wunderlich J. 2006. Fossil and
extant
species of the genus Leptopholcus
in the
Dominican Republic, with the first case of egg-parasitism in pholcid
spiders
(Araneae: Pholcidae). Journal of Natural
History
40: 2341-2360. http://dx.doi.org/10.1111/j.1463-6409.2006.00239.x
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Taczanowski's
pholcids, with three new synonymies in Pholcidae (Araneae). Bull. Br.
arachnol.
Soc. 12(3): 151-152.
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Pérez
González A 1999. The spider genus Crossopriza
(Araneae,
Pholcidae)
in the New World. Amer. Mus. Novitates 3262: 1-10.
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Brescovit
AD, Rheims CA 2005a. Exaggerated female genitalia in
two new
spider species (Araneae: Pholcidae), with comments on genital evolution
by
female choice versus antagonistic coevolution. Insect Syst. Evol. 36:
285-292.
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Pérez González A, Baptista RLC 2005b. Leptopholcus
(Araneae:
Pholcidae)
in continental America: rare relicts in low precipitation areas. Bonner
zool.
Beitr. 53(1/2): 99-107.
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CA, Brescovit AD 2005c. Speciation without changes in
genital
shape: a case study on Brazilian pholcid spiders (Araneae: Pholcidae).
Zool.
Anz. 243: 273-279.
- Huber BA, Rheims
CA, Brescovit AD 2005d. Two new species of litter-dwelling Metagonia
spiders (Araneae, Pholcidae) document both rapid and slow genital
evolution.
Acta Zool. (Stockholm): 86: 33-40.
- Huber BA, Fischer N, Astrin JJ
2010. High level of endemism in Haiti's last remaining forests:
a revision of Modisimus (Araneae:
Pholcidae) on Hispaniola, using morphology and molecules. Zool. J.
Linn. Soc. 158: 244-299.
- Huber BA, Pérez González A, Astrin JJ, Blume C, Baptista R 2013.
Litoporus
iguassuensis
Mello-Leitão, 1918 (Araneae, Pholcidae): camouflaged retreat,
sexual
dimorphism, female color polymorphism, intra-specific genital
variation, and
description of the male. Zoologischer
Anzeiger 252: 511.521.
- Huber BA, Carvalho LS, Benjamin SP 2014a. On
the
New World spiders previously misplaced in Leptopholcus: molecular and
morphological analyses and descriptions of four new species (Araneae,
Pholcidae). Invertebrate
Systematics 28: 432-450.
- Huber BA, Le Gall P, Mavoungou JF
2014b. Pholcid
spiders from the Lower Guinean region of Central Africa: an overview,
with descriptions of seven new species (Araneae, Pholcidae). European
Journal
of Taxonomy 81: 1-46.
- Huber BA, Colmenares PA, Ramirez
MJ 2014c. Fourteen new
generic and ten new specific synonymies in Pholcidae (Araneae), and
transfer
of Mystes Bristowe to Filistatidae. Zootaxa 3847:
413-422.
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BA, Neumann J, Rehfeldt S,
Grabolle A, Reiser N 2015a. Back in Europe: Quamtana spiders (Araneae:
Pholcidsae) in Germany. Arachnologische
Mitteilungen 50: 51-56.
- Huber BA, Nuñeza OM, Leh Moi
Ung
C 2015b. Revision, phylogeny, and microhabitat shifts in the
Southeast Asian spider genus Aetana
(Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78. http://dx.doi.org/10.5852/ejt.2015.162
- Huber BA, Petcharad B, Bumrungsri S
2015c. Revision of the enigmatic Southeast Asian spider genus Savarna (Araneae, Pholcidae). European Journal of Taxonomy 160: 1-23.
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ARM, Nuñeza O, Leh Moi Ung C,
Petcharad B 2016a.
New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia
(Araneae, Pholcidae). European J
Taxonomy 200: 1-45. http://dx.doi.org/10.5852/ejt.2016.200
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BA, Leh Moi Ung C 2016b. The Panjange
nigrifrons group in Borneo (Araneae: Pholcidae): high diversity in
Sarawak,
apparent absence in Sabah. European
Journal of Taxonomy 184: 1-32. http://dx.doi.org/10.5852/ejt.2016.184
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BA,
Nuñeza OM, Leh Moi Ung C 2016c.
The Philippine hair wax spiders and their
relatives: Revision of the Pholcus
bicornutus species group (Araneae, Pholcidae). European
Journal of Taxonomy 225: 1-34.
http://dx.doi.org/10.5852/ejt.2016.225
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BA, Petcharad B, Leh Moi Ung C, Koh JKH, Ghazali ARM 2016d. The Southeast Asian Pholcus halabala species group (Araneae, Pholcidae), new
data from
field observations and ultrastructure. European
Journal of Taxonomy 190: 1-55. http://dx.doi.org/10.5852/ejt.2016.190
- Huber BA, Neumann J, Grabolle A, Hula V.
2017. Aliens in
Europe: updates on the distributions of Modisimus culicinus and Micropholcus fauroti (Araneae, Pholcidae). Arachnologische Mitteilungen 53:
12-18. http://doi.org/10.5431/aramit5303
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BA, Eberle J, Dimitrov D 2018. The
phylogeny of pholcid spiders: a critical evaluation of relationships
suggested by molecular data (Araneae, Pholcidae). ZooKeys 789: 51-101. http://doi.org/10.3897/zookeys.789.22781
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New species reveal
unexpected interspecific microhabitat diversity in the genus Uthina Simon, 1893 (Araneae:
Pholcidae). Invertebrate
Systematics 33: 181-207. https://doi.org/10.1071/IS18002
- Huber BA, Pavlek M, Komnenov M. 2021.
Revision of the spider genus Stygopholcus
(Araneae, Pholcidae), endemic to the Balkan Peninsula. European
Journal of Taxonomy 752: 1-60. https://doi.org/10.5852/ejt.2021.752.1391
- Huber BA, Meng G, Acurio AE, Astrin JJ,
Inclán DJ, Izquierdo M, Valdez-Mondragón A. 2022. Metagonia spiders of
Galápagos: blind cave-dwellers and their epigean relatives
(Araneae, Pholcidae). Invertebrate
Systematics 36(7): 647-678. https://doi.org/10.1071/IS21082
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Herrera M, Inclán D, Wipfler B. 2023a. Humpback spiders
from Ecuador: relationships, prosoma ‘inflation’, and genital asymmetry
(Araneae: Pholcidae: Mecolaesthus).
Invertebrate Systematics 37:
117-151. https://doi.org/10.1071/IS22052
- Huber BA, Meng G, Král J, Ávila Herrera IM, Izquierdo MA, Carvalho LS. 2023b. High and dry: integrative
taxonomy of the Andean spider genus Nerudia
(Araneae: Pholcidae). Zoological
Journal of the Linnean Society 198: 534-591. https://doi.org/10.1093/zoolinnean/zlac100
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Král J, Ávila Herrera IM, Carvalho LS. 2023c. Short-legged daddy-long-leg
spiders in North America: the genera Pholcophora
and Tolteca
(Araneae, Pholcidae). European
Journal of Taxonomy 880: 1-89. https://doi.org/10.5852/ejt.2023.880.2173
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2023d.
First blind daddy long-legs spiders from Australia and Réunion
(Araneae, Pholcidae). Subterranean
Biology 46: 1-19. https://doi.org/10.3897/subtbiol.46.105798
- Huber BA, Meng G, Král J,
Ávila Herrera IM, Izquierdo MA. 2023e. Revision of the
South American Ninetinae genus Guaranita
(Araneae, Pholcidae). European
Journal of Taxonomy 900: 32-80. https://doi.org/10.5852/ejt.2023.880.2173
- Huber BA, Meng G, Dupérré N,
Astrin J, Herrera M. 2023f. Andean giants: Priscula spiders from Ecuador,
with notes on species groups and egg-sac troglomorphism (Araneae:
Pholcidae). European Journal of
Taxonomy 909:1-63. https://doi.org/10.5852/ejt.2023.909.2351
- Huber BA, Meng G, Dederichs TM, Michalik P,
Forman M, Král J. 2024a. Castaways: the Leeward Antilles
endemic spider genus Papiamenta
(Araneae:
Pholcidae). Invertebrate
Systematics 38: IS23052. https://doi.org/10.1071/IS23052
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Valdez-Mondragón A. 2024b. Notes on Chisosa (Araneae: Pholcidae), with
the description of a new species from Mexico. Zootaxa 5419: 217-244. https://doi.org/10.11646/zootaxa.5419.2.3
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García J, Carvalho LS. 2024c. Thriving in dry conditions:
on the Neotropical spider genus Galapa
(Araneae: Pholcidae). Zootaxa
5419: 301-347. https://doi.org/10.11646/zootaxa.5419.3.1
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Progress or burden? Formal description of every apparently new species
available in collections is neither necessary nor useful. ZooKeys 1214:
77-90. https://doi.org./10.3897/zookeys.1214.130592
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